Y-SNP calls for Mathieson (2015) Genome-wide patterns of selection in 230 ancien

Home Miscellanea Population Genomics Y-SNP calls for Mathieson (2015) Genome-wide patterns of selection in 230 ancien

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      Carlos Quiles
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      Mathieson, I., Lazaridis, I., Rohland, N. et al. Genome-wide patterns of selection in 230 ancient Eurasians. Nature 528, 499–503 (2015).

      https://doi.org/10.1038/nature16152

      Abstract:

      Ancient DNA makes it possible to observe natural selection directly by analysing samples from populations before, during and after adaptation events. Here we report a genome-wide scan for selection using ancient DNA, capitalizing on the largest ancient DNA data set yet assembled: 230 West Eurasians who lived between 6500 and 300 BC, including 163 with newly reported data. The new samples include, to our knowledge, the first genome-wide ancient DNA from Anatolian Neolithic farmers, whose genetic material we obtained by extracting from petrous bones, and who we show were members of the population that was the source of Europe’s first farmers. We also report a transect of the steppe region in Samara between 5600 and 300 BC, which allows us to identify admixture into the steppe from at least two external sources. We detect selection at loci associated with diet, pigmentation and immunity, and two independent episodes of selection on height.

      Accession number: PRJEB11450

      Batch Y-chromosome SNP Calls with Yleaf v. 2.3 from FASTQ files, bwa-mem with hg38 reference, and parameters r=1, q=0, b=0.

      Google Drive.

      Haplogroup.info.

      The following is a list of automated Y-DNA haplogroup predictions obtained with Clean tree v2:

      WARNING: Please note that the haplogroup prediction algorithm works best with modern DNA, and its strict requirements make the prediction rather impractical for ancient DNA – in fact, males usually turn out to have not enough quality for a reliable prediction.

      If anything, positive haplogroup predictions (especially under R1b and I2) will normally turn out to be wrong because they are based on few reads, due to the sample being either a) female or b) low coverage.

      The usual process for checking out Y-chromosome haplogroups in ancient DNA samples is as follows:

      1. Select the male individual you want to check out for in the Ancient DNA Dataset.
      2. Look up its library ID (object-ID) at ENA Browser search form (top right) to obtain the accession number(s) for the available sample(s) corresponding to that individual.
      3. Look for files obtained with Yleaf for that accession number(s); more specifically, look for individual Derived and Ancestral SNPs in the ACCESSION_NUMBER.out or ACCESSION_NUMBER.out.csv documents.

      When in doubt about a specific sample, you can ask a question in this thread.

      ERR1136340 NA NA 2305858 1317 0 0 0 0
      ERR1136546 NA NA 8854 3 0 0 0 0
      ERR1136542 NA NA 3439 6 0 0 0 0
      ERR1136393 NA NA 60498 54 0 0 0 0
      ERR1136556 NA NA 4288 7 0 0 0 0
      ERR1136374 NA NA 19739 35 0 0 0 0
      ERR1136337 NA NA 710338 419 0 0 0 0
      ERR1136554 NA NA 8940 9 0 0 0 0
      ERR1136414 NA NA 371508 184 0 0 0 0
      ERR1136391 NA NA 8888583 28682 0 0 0 0
      ERR1136548 NA NA 3002 4 0 0 0 0
      ERR1136401 NA NA 26280 29 0 0 0 0
      ERR1136509 NA NA 1097 4 0 0 0 0
      ERR1136487 R1b1a1b1a1 R-CTS7650*(xFGC14886,YSC0000054,L884.2,S383,S1743,DF27,L356,FGC5538,FGC37092) 1260048 1579 1.0 1.0 1.0 1.0
      ERR1136465 NA NA 3256703 16919 0 0 0 0
      ERR1136400 R1a1 R-M459/etc*(xCTS7083,FGC9998,PF6217,AM00483,KMS2,Y940,Y876,YP4912,Y19836,FGC10223,FGC22471,YP521,YP5906,Y22324,FGC59421,YP1127,S23592,S4576,Y38296,CTS659,YP1506,Y20793,Y34301,YP5579,YP1060,Y23848) 3440082 15893 1.0 1.0 1.0 1.0
      ERR1136434 R1b1a1b1a1a R-PF6543*(xAM01884,A561,A1243,A680,S5556,S24013,FGC12316,JFS0008,FGC6184,Z159,S515,A9456,FGC17362,FGC19771,S15510,S1731,CTS8234,S19882,S23465,PF7595,L276,BY129,L617,L881,F3867.2,DF81,S370,S20782,FGC31475,PF6584,L583,A7907,Z2245,ZZ38) 1313932 4647 1.0 1.0 1.0 1.0
      ERR1136397 R1a1a1b2a~ R-F3568*(xAM00483,YP4912,FGC10223,FGC22471,Y36348,Y34161,YP5279,YP4886,YP1547,Y12480,F1345,YP641,FGC4547,CTS659) 1466752 6398 1.0 1.0 1.0 1.0
      ERR1136339 NA NA 404693 273 0.0 0.0 1.0 1
      ERR1136344 NA NA 97250 85 0 0 0 0
      ERR1136423 NA NA 42377 57 0 0 0 0
      ERR1136428 NA NA 6660929 24023 0.0 0.0 1.0 0.0
      ERR1136463 R1b1a1b1a1 R-CTS7650*(xL884.2,FGC915,S1746,S22641,DF27,L356,L564,L644,Z44612) 669387 666 1.0 1.0 1.0 1.0
      ERR1136360 R1b1a2 R-BY15360/etc*(xBY15332,Y109041) 1985109 8822 1.0 1.0 1.0 1.0
      ERR1136460 NA NA 3280985 17328 0 0 0 0
      ERR1136431 T1a T-M70*(xCTS8512) 572876 1531 1.0 1.0 1.0 1.0
      ERR1136351 NA NA 598624 310 0 0 0 0
      ERR1136550 NA NA 32957 57 0 0 0 0
      ERR1136437 NA NA 954265 2115 0 0 0 0
      ERR1136425 NA NA 358659 648 0 0 0 0
      ERR1136406 R1b1a1b R-PF6433/etc*(xL51,CTS9219,GG644,FGC42031) 1100561 3851 1.0 1.0 1.0 1.0
      ERR1136447 NA NA 36971 38 0 0 0 0
      ERR1136482 NA NA 35613 16 0 0 0 0
      ERR1136526 NA NA 3745 3 0 0 0 0
      ERR1136356 NA NA 275129 188 0 0 0 0
      ERR1136504 NA NA 27798 40 0 0 0 0
      ERR1136408 NA NA 43806 46 0 0 0 0
      ERR1136421 NA NA 1379577 5912 0.0 0.0 1.0 1.0
      ERR1136523 NA NA 14039 12 0.0 0.0 1.0 0.0
      ERR1136382 R1b1a1b R-CTS11468/etc*(xPF6540,PF7589,Z2106,GG480) 2200090 9908 1.0 1.0 1.0 1.0
      ERR1136483 NA NA 461058 286 0 0 0 0
      ERR1136541 NA NA 8025 14 0 0 0 0
      ERR1136461 NA NA 3236861 16837 0 0 0 0
      ERR1136470 NA NA 3098414 16416 0 0 0 0
      ERR1136507 NA NA 5892 5 0 0 0 0
      ERR1136384 H2a H-Z19120*(xZ18952) 822882 3626 1.0 1.0 1.0 1.0
      ERR1136398 R1a1a1b2 R-F992*(xAM00483,YP4912,Y19762,FGC10223,FGC22486,YP521,YP5277,M12291,YP1561,YP1353,S18885,M12202,FGC131,YP5959,PH4192,Y57,FGC30901,CTS659,Y34301,YP5591) 1826230 7629 1.0 1.0 1.0 1.0
      ERR1136417 NA NA 3369810 10406 0 0 0 0
      ERR1136486 R1b1a1b R-CTS1416^^/etc*(xFGC36479,L1231,L356,L564,FGC31939) 1797708 2007 1.0 1.0 1.0 1.0
      ERR1136479 R1a1a1~ R-F3159/etc*(xYP5531,YP5318,YP367,YP442,YP4515,FGC10010,F3044) 666768 3549 1.0 1.0 1.0 1.0
      ERR1136450 NA NA 55602 77 0 0 0 0
      ERR1136543 NA NA 759 2 0 0 0 0
      ERR1136375 NA NA 498509 329 0 0 0 0
      ERR1136372 NA NA 269635 262 0 0 0 0
      ERR1136565 NA NA 7566 2 0 0 0 0
      ERR1136331 I2a1a1b~ I-S15399*(xL1286,A10111) 1471132 8754 1.0 1.0 1.0 1.0
      ERR1136379 R1a1 R-M459*(xYP368,FGC10010,BY32079,AMM264,YP569,Y1400,FGC19244,FGC21243,YP6086,Y33119,YP1628,YP1148,CTS8277,YP6455,Y10817,Y16851,YP1370,Y34256,Y19831,FGC10205,Y36346,YP5277,YP1557,Y37891,FGC30903,Y23848) 218209 746 1.0 1.0 1.0 1.0
      ERR1136512 NA NA 52085 66 0 0 0 0
      ERR1136352 NA NA 370752 217 0.0 0.0 1.0 1.0
      ERR1136469 NA NA 494057 536 0 0 0 0
      ERR1136433 I2a2 I-PF3897*(xY16665,Y91571) 387384 850 1.0 1.0 1.0 1.0
      ERR1136390 R1 R-L875*(xM768,PF6463,BY21441,V35,BY17639) 313369 829 1.0 1.0 1.0 1.0
      ERR1136330 NA NA 343040 252 0 0 0 0
      ERR1136413 R1b1 R-L761/etc*(xCTS11985,PF6314) 609905 2211 1.0 1.0 1.0 1.0
      ERR1136388 NA NA 372756 221 0 0 0 0
      ERR1136430 NA NA 770802 2122 0 0 0 0
      ERR1136418 R1b1a1b1a1 R-CTS7650*(xAM01880,YSC0000054,S263,FGC419,L1231,L356,S653,CTS2501,S470,L743,S279,CTS8358,DF73,L1308,S3106,S281,L130,FGC58892,FGC33372,L1315,BY11894,FGC37092) 1768846 1417 1.0 1.0 1.0 1.0
      ERR1136510 NA NA 19793 13 0 0 0 0
      ERR1136442 NA NA 1377798 7943 0 0 0 0
      ERR1136380 NA NA 1450575 5563 0.0 0.0 1.0 1.0
      ERR1136335 NA NA 812249 763 0 0 0 0
      ERR1136396 NA NA 478979 381 0 0 0 0
      ERR1136377 NA NA 125841 112 0 0 0 0
      ERR1136501 NA NA 31674 47 0 0 0 0
      ERR1136452 NA NA 71929 105 0 0 0 0
      ERR1136399 R1b1a1b1b R-Z2105*(xCTS7822) 2023747 9140 1.0 1.0 1.0 1.0
      ERR1136522 NA NA 6381 14 0 0 0 0
      ERR1136540 NA NA 6725 6 0 0 0 0
      ERR1136462 NA NA 642325 685 0.0 1.0 1.0 0.0
      ERR1136558 NA NA 25496 39 0 0 0 0
      ERR1136506 NA NA 72067 93 0 0 0 0
      ERR1136386 NA NA 394146 298 0 0 0 0
      ERR1136473 NA NA 1243683 6733 0.0 0.0 1.0 1.0
      ERR1136395 NA NA 45686 42 0 0 0 0
      ERR1136424 R1b1a1b R-CTS1416^^/etc*(xL884.2,S1743,L356,BY281,Z18064.1,BY404,S280,FGC33060,BY35673,AM01878) 215378 290 1.0 1.0 1.0 1.0
      ERR1136513 NA NA 3946 5 0 0 0 0
      ERR1136348 NA NA 3038167 2248 0.0 0.0 1.0 1.0
      ERR1136488 NA NA 6403844 24941 0 0 0 0
      ERR1136531 NA NA 4226 3 0 0 0 0
      ERR1136459 G2a G-PF3141*(xFGC621^,PF3181,F2184) 480279 3373 1.0 1.0 1.0 1.0
      ERR1136383 NA NA 3103223 15825 0.0 0.0 1.0 1.0
      ERR1136484 NA NA 225610 130 0 0 0 0
      ERR1136359 R1b1a1b1 R-L23*(xS263,L1231,L356,L192.1,FGC37092) 2076052 1267 1.0 1.0 1.0 1.0
      ERR1136464 R1b1a1b1 R-L23*(xS263,L1231,L356,L459) 651800 694 1.0 1.0 1.0 1.0
      ERR1136458 NA NA 848061 6648 0.0 0.0 1.0 1.0
      ERR1136376 NA NA 609960 440 0.0 0.0 1.0 1.0
      ERR1136405 R1b1a1b1b3a R-CTS1843*(xS12460) 1916065 8443 1.0 1.0 1.0 1.0
      ERR1136346 NA NA 5229443 21002 0.0 0.0 1.0 1.0
      ERR1136364 NA NA 49285 38 0 0 0 0
      ERR1136511 NA NA 11106 8 0 0 0 0
      ERR1136477 NA NA 33725 21 0 0 0 0
      ERR1136525 NA NA 7940 9 0 0 0 0
      ERR1136537 NA NA 3944 3 0 0 0 0
      ERR1136485 NA NA 1018160 1294 0 0 0 0
      ERR1136410 R1b1a1b1b R-CTS1078*(xL277.1) 1783583 7777 1.0 1.0 1.0 1.0
      ERR1136367 NA NA 81623 45 0 0 0 0
      ERR1136353 NA NA 885567 351 0 0 0 0
      ERR1136476 R1a1a1~ R-CTS10993/etc*(xYP548,YP6488,FGC10012,CTS9754) 311915 1669 1.0 1.0 1.0 1.0
      ERR1136481 R1a1a1~ R-CTS10993*(xY4393,CTS9754) 278756 1477 1.0 1.0 1.0 1.0
      ERR1136389 NA NA 2049240 7701 0.0 1.0 1.0 0.0
      ERR1136347 R1b1a1b1 R-L23*(xFGC14886,A564,YSC0000054,S263,FGC390,L1231,L737,Z49,FGC22528,L459,AM01878,A8053,PF7585) 1635205 1275 1.0 1.0 1.0 1.0
      ERR1136407 R1b1a1b1 R-L23*(xL51,PF7585) 6498812 25233 1.0 1.0 1.0 1.0
      ERR1136361 NA NA 1333202 6004 0.0 0.0 1.0 1.0
      ERR1136329 I2a1a2 I-M423/etc*(xL161.1,S19848) 328593 2202 1.0 1.0 1.0 1.0
      ERR1136336 R1b1a1b R-CTS329/etc*(xFGC30616,FGC13260,S1743,L1231,Z49,L192.1) 934850 400 1.0 1.0 1.0 1.0
      ERR1136332 NA NA 4746931 19951 0 0 0 0
      ERR1136534 NA NA 254 2 0 0 0 0
      ERR1136478 R1a1a1~ R-M630/etc*(xYP5528,Z2035,YP5318,FGC19043,Y34274,YP4101,FGC9997,F3044) 659942 3926 1.0 1.0 1.0 1.0
      ERR1136503 NA NA 5139 6 0 0 0 0
      ERR1136508 NA NA 15143 18 0 0 0 0
      ERR1136555 NA NA 40176 33 0 0 0 0
      ERR1136350 NA NA 105668 75 0 0 0 0
      ERR1136441 R1b1a1b R-L150.1*(xYSC0000054,S1743,S22641,L1231,L356,S653,A11313,Z17836,Z21255,BY404,BY2743,BY357,S5456,CTS8704,FGC37083) 495422 595 1.0 1.0 1.0 1.0
      ERR1136333 NA NA 213436 228 0 0 0 0
      ERR1136467 NA NA 1780472 11553 0.0 0.0 1.0 1.0
      ERR1136371 NA NA 304859 243 0.0 1.0 1.0 0.0
      ERR1136378 NA NA 436343 282 0 0 0 0
      ERR1136530 NA NA 27072 20 0 0 0 0
      ERR1136358 NA NA 1093040 678 0 0 0 0
      ERR1136369 NA NA 8438001 29729 0.0 0.0 1.0 1.0
      ERR1136572 NA NA 7304 2 0 0 0 0
      ERR1136445 NA NA 948243 5671 0 0 0 0
      ERR1136438 R1b1a1b R-CTS1415^^/etc*(xYSC0000054,Y10790,Y7369,S1746,L1231,L356,S6154,L746,FGC5606,Z16351,BY404,A1334,FGC5654,S844,BY357,S5456,FGC43487,FGC33060,FGC58892,ZZ38) 754181 865 1.0 1.0 1.0 1.0
      ERR1136520 NA NA 86912 87 0 0 0 0
      ERR1136357 NA NA 2724715 10768 0 0 0 0
      ERR1136527 NA NA 5187 2 0 0 0 0
      ERR1136457 NA NA 117466 773 0.0 0.0 1.0 0.0
      ERR1136567 NA NA 8467 14 0 0 0 0
      ERR1136394 R1a~ R-CTS10627/etc*(xY177,YP4145) 2130195 7773 1.0 1.0 1.0 1.0
      ERR1136471 R1b1a1b1a1 R-CTS7650*(xS499,DF79,L356,S653,FGC5606,L743,Z17836,BY404,FGC58883,L1315,AM01878,FGC37092) 675121 888 1.0 1.0 1.0 1.0
      ERR1136404 Q1 Q-F2676*(xF4930,CTS950) 143257 599 1.0 1.0 1.0 1.0
      ERR1136370 NA NA 10018854 34050 0.0 0.0 1.0 1.0
      ERR1136327 NA NA 330589 204 0 0 0 0
      ERR1136551 NA NA 14296 7 0 0 0 0
      ERR1136366 H2 H-Z18831/etc*(xZ19110,Z41305,Y21618) 876492 2790 1.0 1.0 1.0 1.0
      ERR1136436 NA NA 742661 1918 0.0 0.0 1.0 0.0
      ERR1136342 NA NA 75101 64 0 0 0 0
      ERR1136518 NA NA 26796 7 0 0 0 0
      ERR1136480 R1b1a2 R-CTS6666 227721 1036 1.0 1.0 1.0 1.0
      ERR1136368 NA NA 391525 1844 0.0 1.0 1.0 0.0
      ERR1136453 NA NA 259002 350 0 0 0 0
      ERR1136354 NA NA 449577 251 0 0 0 0
      ERR1136432 NA NA 345354 307 0 0 0 0
      ERR1136547 NA NA 2901 4 0 0 0 0
      ERR1136328 NA NA 3075862 13992 0 0 0 0
      ERR1136455 I2a I-CTS4039*(xCTS6406,CTS12005,Z27379,Y19490,M423,FGC14219,Y7243,Y4904,Y5383,Y7494,P95,Y22356,Y24165,Y7831,Y5717,Y13439,FGC20016,BY3605,FGC30172,Y31078,Y36058,Y36558,Y15494,Y5190,BY44566,Y18922,L597) 176895 1374 1.0 1.0 1.0 1.0
      ERR1136427 NA NA 3970177 16417 0 0 0 0
      ERR1136446 NA NA 1584675 9468 0 0 0 0
      ERR1136402 R1a1a1b2a~ R-F3568*(xAM00479,Y18102,YP4912,Y16006,YP4976,Y19762,FGC10207,FGC22471,Y36346,YP5906,FGC23228,Y22323,FGC59402,YP1709,S23592,S4576,FGC30903,Z664) 2395157 10438 1.0 1.0 1.0 1.0
      ERR1136561 NA NA 519 1 0 0 0 0
      ERR1136475 R1a R-L62*(xYP5534,FGC32020,AMM259,A8993,BY30793,FGC19232,YP4835,Y17527,YP4820,Y10895,Y19886,FGC10190,Y16501,L448,F19106,S6351,YP409,FGC28678,FGC64140,Y32864,M12369,FGC73292,Y2447,Y609,Y36351,FGC23228,FGC48776,YP1300,Y22243) 276499 1508 1.0 1.0 1.0 1.0
      ERR1136422 I2a2 I-PF3893*(xFGC18609) 150489 636 1.0 1.0 1.0 1.0
      ERR1136343 NA NA 438706 340 0 0 0 0
      ERR1136355 NA NA 3291053 13165 0.0 0.0 1.0 1.0
      ERR1136449 I2a1b1b1a~ I-S24785 218358 1855 1.0 1.0 1.0 1.0
      ERR1136392 NA NA 474104 413 0 0 0 0
      ERR1136341 NA NA 883756 4203 0 0 0 0
      ERR1136568 NA NA 6557 7 0 0 0 0
      ERR1136474 NA NA 338047 2036 0.0 1.0 1.0 0.0ERR1136444 NA NA 1886019 11119 0.0 0.0 1.0 0.0
      
      ERR1136566 NA NA 4222 5 0 0 0 0
      ERR1136529 NA NA 3710 1 0 0 0 0
      ERR1136409 NA NA 5901155 23660 0.0 0.0 1.0 1.0
      ERR1136387 R1b1a1b1a1 R-CTS7650*(xYSC0000054,S263,L1231,L356,L192.1,Z2248,A8053) 2360006 2262 1.0 1.0 1.0 1.0
      ERR1136535 NA NA 3008 4 0 0 0 0
      ERR1136549 NA NA 9189 13 0 0 0 0
      ERR1136411 NA NA 198083 109 0 0 0 0
      ERR1136489 R1b1a1b1a1a2c1a5b1 R-CTS8704*(xFGC5783) 1135720 1276 1.0 1.0 1.0 1.0
      ERR1136468 R1b1a1b R-PF6462/etc*(xS263,L1231,L356,FGC19851,BY281,L743,Z18064.1,S844,FGC5782,FGC43488,FGC33060,FGC42968,L1315,Z2248,AM01878) 610980 682 1.0 1.0 1.0 1.0
      ERR1136338 NA NA 1607603 6355 0.0 0.0 1.0 1.0
      ERR1136419 NA NA 5940344 21801 0 0 0 0
      ERR1136403 R1a1 R-M459*(xM515,YP1335) 1431179 6077 1.0 1.0 1.0 1.0
      ERR1136362 NA NA 1272967 4581 0.0 0.0 1.0 1.0
      ERR1136420 NA NA 6297418 23262 0.0 0.0 1.0 0.0
      ERR1136563 NA NA 3567 4 0 0 0 0
      ERR1136439 NA NA 2212851 13455 0.0 0.0 1.0 1.0
      ERR1136363 NA NA 547098 279 0 0 0 0
      ERR1136334 R1b1a1b1a1a1c R-S263*(xS384,S1743,S22641) 825257 770 1.0 1.0 1.0 1.0
      ERR1136552 NA NA 453 2 0 0 0 0
      ERR1136451 NA NA 1094826 8275 0 0 0 0
      ERR1136440 NA NA 2161932 13185 0 0 0 0
      ERR1136429 NA NA 6439066 23995 0 0 0 0
      ERR1136559 NA NA 37138 39 0 0 0 0
      ERR1136415 NA NA 7189369 26611 0 0 0 0
      ERR1136426 NA NA 712674 628 0 0 0 0
      ERR1136557 NA NA 366 3 0 0 0 0
      ERR1136456 R1b1a1b1 R-L23*(xFGC36479,L1231,L356,L564,CTS300) 517998 580 1.0 1.0 1.0 1.0
      ERR1136528 NA NA 64408 50 0 0 0 0
      ERR1136416 NA NA 9511735 30110 0 0 0 0
      ERR1136412 G2a2a1 G-S11510*(xFGC6691,PF3233,FGC34624,FGC34447) 664138 1707 1.0 1.0 1.0 1.0
      ERR1136502 NA NA 18053 27 0 0 0 0
      ERR1136533 NA NA 1172 3 0 0 0 0
      ERR1136544 NA NA 660 2 0 0 0 0
      ERR1136373 NA NA 193083 1008 0 0 0 0
      ERR1136539 NA NA 370 2 0 0 0 0
      ERR1136443 NA NA 149350 251 0 0 0 0
      ERR1136381 NA NA 2922063 12386 0.0 0.0 1.0 1.0
      ERR1136466 NA NA 691401 641 0 0 0 0
      ERR1136472 R1b1a1b R-CTS623*(xA1243,A7409,S385,S12025,Z272,Z203,L881,F1493.2,A1332,CTS2501,Z16330,Z17830,Z23534,A6507,BY3455,BY19857,Z2247,DF99,GG644,Y31469) 66445 417 1.0 1.0 1.0 1.0
      ERR1136365 NA NA 13200716 39189 0 0 0 0
      ERR1136521 NA NA 6031 12 0 0 0 0
      ERR1136435 R1b1a1b1a1a2 R-P312*(xDF17,BY132,A431,L409,CTS4333,BY247,Z2248,ZZ38) 939695 2907 1.0 1.0 1.0 1.0
      ERR1136345 R1a1 R-M459/etc*(xM514,Y12636) 5766288 23924 1.0 1.0 1.0 1.0
      ERR1136349 NA NA 5452525 22202 0.0 0.0 1.0 1.0
      ERR1136448 NA NA 164063 163 0 0 0 0
      ERR1136569 NA NA 12845 24 0 0 0 0
      ERR1136385 NA NA 2771358 12028 0 0 0 0
      ERR1136514 NA NA 2238 5 0 0 0 0
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